Arbeitsgruppe Morphologie und Systematik der Phanerogamen

Boraginales (Boraginaceae s.l.) - History -

Hilger, H.H., Cole, T.H.C. & Selvi, F. (2014): J.G.C. Lehmann's 'Botanical Observations' of 1818 on Coldenia, Colsmannia, Cynoglossum, and Omphalodes (commented translation). - Biodiv. Data J. 2: e1064. http://dx.doi.org/10.3897/BDJ.2.e1064.

Boraginales (Boraginaceae s.l.) - general -

Luebert, F.; Couvreur, T.L.P., Gottschling, M., Hilger, H.H., Miller, J.S., Weigend, M. (2017): Historical biogeography of Boraginales: West Gondwanan vicariance followed by long-distance dispersal?. - J. Biogeogr. 44: 158-169.
Luebert, F., Cecchi, L., W. Frohlich, M.W., Gottschling, M., Guilliams, C.M., Hasenstab-Lehman, H.E., Hilger, H.H., Miller, J.S., Mittelbach, M., Nazaire, M., Nepi, M., Nocentini, D., Ober, D., Olmstead, R.G., Selvi, F., Simpson, M.G., Sutorý, K., Valdés, B., Walden G.K. & Weigend, M. (2016): Familial classification of the Boraginales. - Taxon 65(3): 502-522
Weigend, M., Luebert, F., Gottschling, M., Couvreur, T.L.P., Hilger, H.H. & Miller, J.S. (2014): From capsules to nutlets — phylogenetic relationships in the Boraginales. - Cladistics 30(5): 508-518. DOI: 10.1111/cla.12061
Hilger, H.H., Selvi, F., Bigazzi, M., Langstrom, E., Gottschling, M., Zippel, E., Diane, N. & Weigend, M. (2005): The Euro+Med treatment of Boraginaceae - a response. - Willdenowia 35 (1): 43-48.
Gottschling, M., Diane, N., Hilger, H.H. & Weigend, M. (2004): Testing hypotheses on disjunctions present in the Primarily Woody Boraginales: Ehretiaceae, Cordiaceae, and Heliotropiaceae, inferred from ITS1 sequence data. - Int. J. Plant Sci. 165 (4 Suppl.): S123-S135.
Hilger, H.H. & Förther, H. (2004): „Boraginaceae“ - in: Neotropical Plant Families - ed. New York Botanical Garden.
Hofmann, M. (2004): "Hydrophyllaceae" - in: Neotropical Plant Families - ed. New York Botanical Garden
Buys, M.H. & Hilger, H.H. (2003): Boraginaceae cymes are exclusively scorpioid and not helicoid. - Taxon 52: 719-724.
Diane, N., Hilger, H.H. & Gottschling, M. (2002): Transfer cells in the seeds of Boraginales . - Bot. J. Linn. Soc. 140: 155-164.
Gottschling, M., Hilger, H.H., Wolf, M. & Diane, N. (2001): Secondary Structure of the ITS1 Transcript and its Application in a Reconstruction of the Phylogeny of Boraginales. - Plant biol. 3: 629-636.
Hilger, H.H. & Zippel, E. (2001): "Studies in the Boraginaceae" - An Index to the Publications of Ivan M. Johnston dealing with the Borage Family. - Haussknechtia Beiheft 11: 1-151. Jena.
Briechle, M. & Hilger, H.H. (1988): Die Embryogenese von Microparacaryum intermedium und die embryologische Klassifizierung der Boraginaceae nach Souèges. - Flora 181: 45-59.

Boraginaceae s.str.

Weigend, M., Selvi, F., Thomas, D. & Hilger, H.H. (2016): Boraginaceae. pp. 41-102 in Kadereit, J.W. & Bittrich, V. (eds.): The Families and Genera of Vascular Plants Vol. 14: Flowering Plants. Eudicots. Aquifoliales, Boraginales, Bruniales, Dipsacales, Escalloniales, Garryales, Paracryphiales, Solanales (except Convolvulaceae), Icacinaceae, Metteniusaceae, Vahliaceae. Springer - Switzerland.
Chacón, J., Luebert, F., Hilger; H.H., Ovchinnikova, S., Selvi, F., Cecchi, L., Guilliams, C.M., Hasenstab-Lehman, K., Sutorý, K., Simpson, M.G., Weigend, M. (2016): The borage family (Boraginaceae s.s.): A revised infrafamilial classification based on new phylogenetic evidence, with emphasis on the placement of some enigmatic genera. - Taxon 65(3): 523-546
Weigend, M., Luebert, F., Selvi, F., Brokamp, G. & H.H. Hilger (2013): Multiple origins for Hounds tongues (Cynoglossum L.) and Navel seeds (Omphalodes Mill.) – the phylogeny of the borage family (Boraginaceae s.str.). - Molecular Phylogenetics and Evolution 68(3): 604-618.
Hilger, H.H., Hoppe, J.R. & Hofmann, M. (1993): Energiedispersive Röntgenmikroanalyse (EDX) von Boraginaceae subfam. Boraginoideae-Klausenoberflächen (Sind Silicium- und Calcium-Einlagerungen in die Fruchtwand systematisch verwertbare Merkmale?). - Flora 188: 387-398.

Lithospermeae

Muller, L.A.H. & H.H. Hilger (2015): Insights into the effects of serpentine soil conditions on the community composition of fungal symbionts in the roots of Onosma echioides. - Soil Biol. Biochem. 81: http://www.sciencedirect.com/science/article/pii/S0038071714003745
Cecchi, L., A. Coppi, H.H. Hilger & F. Selvi (2014): Non-monophyly of Buglossoides (Boraginaceae: Lithospermeae): Phylogenetic and morphological evidence for the expansion of Glandora and reappraisal of Aegonychon. - Taxon 63: 1965-1078
Weigend, M., Gottschling, M., Hilger, H.H. & Nürk, N.M. (2010): Five new species of Lithospermum L. (Boraginaceae tribe Lithospermeae) in Andean South America - another radiation in the Amotape-Huancabamba Zone. - Taxon 59(4): 1161-1179.
Pustovoytov, K., Riehl, S., Hilger, H.H. & Schumacher, E. (2010): Oxygen isotopic composition of fruit carbonate in Lithospermeae and its potential for paleoclimate research in the Mediterranean. - Global Planet. Change 71: 258-268.
Weigend, M., Gottschling, M., Selvi, F. & Hilger, H.H. (2009): Marbleseeds are gromwells - Systematics and evolution of Lithospermum and allies (Boraginaceae tribe Lithospermeae) based on molecular and morphological data. - Molec. Phylogen. Evol. 52 (3): 755-768.
Thomas, D.C., Weigend, M. & Hilger, H.H. (2008): Phylogeny and systematics of Lithodora (Boraginaceae, Lithospermeae) and its affinities to the monotypic genera Neatostema, Mairetis, Halacsya and Paramoltkia based on ITS1 and trnLUAA-sequence data and morphology. - Taxon 57 (1): 79-97.
Cusimano, N. (2008 publ. 2010): Onosma echioides L. (Boraginaceae) besiedelt Serpentinböden in der Toskana (Italien). - Sber. Ges. Naturf. Freunde Berlin (N.F.) 47: 93-101.
Mengoni, A., Selvi, F., Cusimano, N., Galardi, F., Gonnelli, C. (2006): Genetic diversity inferred from AFLP fingerprinting in populations of Onosma echioides (Boraginaceae) from serpentine and calcareous soils. - Plant Biosystems 140 (2): 211-219.
Clermont, A., Zippel, E. & Hilger, H.H. (2003): Verbreitung und Differenzierung der mitteleuropäischen Unterarten von Buglossoides arvensis (L.) I.M.Johnst.(Boraginaceae). - Feddes Repert. 114 (1-2): 58-70.
Hilger, H.H. & Böhle, U.-R. (2000): Pontechium - a new genus separated from Echium and Lobostemon (Boraginaceae). - Taxon 49: 737-746.
Böhle, U.-R., Hilger, H.H. & Martin, W. (1996): Island colonization and evolution of the insular woody habit in Echium L. (Boraginaceae). - Proc. Natl. Acad. Sci. USA 93: 11740-11745.
Hilger, H.H. (1994): Die Evolution der Rauhblattgewächse (Boraginaceae) in Makaronesien. - Ulmensien 8: 23-32.
Böhle, U.-R., Hilger, H.H., Cerff, R. & Martin, W. (1994): Non-coding chloroplast DNA for plant molecular systematics at the infrageneric level. - pp. 391-403 in: Schierwater, B., Streit, B., Wagner, G.P. & DeSalle, R. [eds.], Molecular Ecology and Evolution: Approaches and Application. Birkhäuser, Basel.


Boragineae

Weigend, M., Gottschling, M., Selvi, F. & Hilger, H.H. (2010): Fossil and extant Western hemisphere Boragineae, and the polyphyly of "Trigonotideae" Riedl (Boraginaceae: Boraginoideae). - Syst. Bot. 35(2): 409-419.
Selvi, F., Bigazzi, M., Hilger, H.H., & Papini, A. (2006): Molecular phylogeny, morphology and taxonomic re-circumscription of the generic complex Nonea/Elizaldia/Pulmonaria/Paraskevia (Boraginaceae - Boragineae). - Taxon 55 (4): 907-918.
Bigazzi, M., Selvi, F. & Hilger, H.H. (2004): Nonea pisidica (Boraginaceae-Boragineae), a new species from southwest Anatolia and its relationships inferred from karyology and cpDNA sequences. - Plant Biosystems 138 (2): 135-144.
Hilger, H.H., Selvi, F., Papini, A. & Bigazzi, M. (2004): Molecular systematics of Boraginaceae tribe Boragineae based on ITS1 and trnL sequences, with special reference to Anchusa s.l. - Ann. Bot. 94 (2): 201-212.
Selvi, F., Papini, A., Hilger, H.H., Bigazzi, M. & Nardi, E. (2004): The phylogenetic relationships of Cynoglottis (Boraginaceae-Boragineae) inferred from ITS, 5.8S and trnL sequence variation - Pl. Syst. Evol. 246 (3-4): 195-209.
Bigazzi, M., Hilger, H,H. & Selvi, F. (2002): Evidence from nuclear and chloroplast DNA for the placement of Anchusa macedonica in the genus Gastrocotyle (Boraginaceae).- Webbia 57 (2): 173-180.
Danin, A. (2001): Hormuzakia negenvensis (Danin) Danin & Hilger comb. nov., pp. 110-111 in Danin, A.: The nomenclature news of Flora Palaestina. - Fl. Medit. 10: 109-172.


Cynoglosseae [incl. Eritrichieae]

Holstein, N., Chacón, J., Hilger, H.H. & Weigend, M. (2016): No longer shipwrecked – Selkirkia (Boraginaceae) back on the mainland with generic rearrangements in South American Omphalodes based on molecular data. - Phytotaxa 270(4): 231-251.
Hilger, H.H.: Greuter, W. & Stier, V. (2015): Taxa and names in Cynoglossum sensu lato (Boraginaceae, Cynoglosseae): an annotated, synonymic inventory, with links to the protologues and mention of original material. - Biodiversity Data Journal 3: e4831 (22 Apr 2015) doi: 10.3897/BDJ.3.e4831.
König, J., Siwakoti, M., Hilger, H.H. & Weigend, M. (2015): A revision of the genus Cynoglossum L. (Boraginaceae Juss.) in Nepal and notes on the widespread Asian species. - Phytotaxa 224(1): 1-28. http://biotaxa.org/Phytotaxa/article/view/phytotaxa.224.1.1/18191
Dickoré, W.B. & Hilger, H.H. (2015): Decalepidanthus (Boraginaceae) includes and antedates Pseudomertensia; a synopsis of the genus. - Phytotaxa 226(2): 131-143.
Greuter, W., Stier, V. & Hilger, H.H. (2014): Proposal to conserve the name Omphalodes verna against Omphalodes omphaloides (Boraginaceae). - Taxon 63 (2): 435.
Hilger, H.H. (2014): Ontogeny, morphology, and systematic significance of glochidiate and winged fruits of Cynoglosseae and Eritrichieae (Boraginaceae). - Pl. Div. Evol. 131(3): 167-214.
Hofmann, M. (1994): Untersuchungen zur Gynoecealentwicklung in der Tribus Eritrichieae (Boraginaceae-Boraginoideae). Doct. thesis FU Berlin, 311 pp. UB 88/95/60193(9)
Hilger, H.H. (1986): Mikromorphologische Untersuchungen an Klausenoberflächen von Boraginaceae II. Mattiastrum (Boiss.) Brand (Cynoglosseae). - Beiträge zur Biologie der Pflanzen 61: 145-165.
Hilger, H.H. (1985): Ontogenie, Morphologie und systematische Bedeutung geflügelter und glochidientragender Cynoglosseae- und Eritrichieae-Früchte (Boraginaceae). - Botanische Jahrbücher für Systematik, Pflanzengeschichte und Pflanzengeographie 105: 323-378.
Hilger, H.H., Balzer, M. & Podlech, D. (1984): Heteromerikarpie und Fruchtpolymorphismus bei Microparacaryum, gen. nov. (Boraginaceae). - Plant Systematics and Evolution 148: 291-312.
Hilger, H.H. (2014): Wachstum und Ausbildungsformen des Gynoeceums von Rochelia (Boraginaceae). - Plant Systematics and Evolution 146: 123-139.
Hilger, H.H. & Richter, U. (1982): Untersuchungen zur Ausbildung der Klausen und ihrer Oberflächenskulpturen bei Paracaryum intermedium (Boraginaceae) aus Jordanien. - Beiträge zur Biologie der Pflanzen 57: 205-220.
Hilger, H.H. (1981): Entwicklungstendenzen im Fruchtbau der anemochoren Boraginaceengattungen Mattiastrum und Paracaryum. - Berichte der Deutschen Botanischen Gesellschaft 94: 677-687.
Hilger, H.H. (1981): Die Perikarpentwicklung geflügelter Klausen aus dem Paracaryum - Verwandtschaftskreis (Cynoglosseae, Boraginaceae). - Plant Systematics and Evolution 138: 73-88.

Cordiaceae und Ehretiaceae

Gottschling, M., Weigend, M. & Hilger, H.H. (2016): Ehretiaceae incl. Cordiaceae. pp. 165-178 in Kadereit, J.W. & Bittrich, V. (eds.): The Families and Genera of Vascular Plants Vol. 14: Flowering Plants. Eudicots. Aquifoliales, Boraginales, Bruniales, Dipsacales, Escalloniales, Garryales, Paracryphiales, Solanales (except Convolvulaceae), Icacinaceae, Metteniusaceae, Vahliaceae. Springer - Switzerland.
Gottschling, M., Nagelmüller, S. & Hilger, H.H. (2014): Generative ontogeny in Tiquilia (Ehretiaceae, Boraginales) and phylogenetic implications. - Biol. J. Linn. Soc. 112: 520-534
Gottschling, M., Luebert, F., Hilger, H.H. & Miller, J.S. (2014): Molecular delimitations in the Ehretiaceae (Boraginales). - Mol. Phylogenet. Evol. DOI:10.1016/j.ympev.2013.12.005
Gottschling, M. & Miller, J.S. (2007): A revision of Bourreria (Ehretiaceae, Boraginales) in South America. - Ann. Mo. Bot. Gard. 94 (4): 734-744
Gottschling, M. & Miller, J.S. (2007): Typification of Bourreria names (Ehretiaceae, Boraginales) based on specimens collected by Charles Wright in Cuba. - Taxon 56 (1): 237-242.
Miller, J.S. & Gottschling, M. (2007): Generic classification in the Cordiaceae (Boraginales): resurrection of the genus Varronia P.Br.- Taxon 56 (1): 163-169.
Gottschling, M. & Miller, J.S. (2006): Clarification of the taxonomic position of Auxemma, Patagonula, and Saccellium (Cordiaceae, Boraginales). - Syst. Bot. 31 (2): 361-367.
Gottschling, M., Miller, J.S., Weigend, M. & Hilger, H.H. (2005): Congruence of a phylogeny of Cordiaceae (Boraginales) inferred from ITS1 sequence data with morphology, ecology, and biogeography. - Ann. Missouri Bot. Gard. 92 (3): 425-437. Gottschling, M. & Miller, J.S. (2005): A new species of Bourreria (Ehretiaceae, Boraginales) from Costa Rica. - Novon 15: 425-428.
Gottschling, M. & Hilger, H.H. (2004): The systematic position of Ehretia cortesia nom. nov. (syn. Cortesia cuneifolia:Ehretiaceae, Boraginales) inferred from molecular and morphological data. - Taxon 53 (4): 919-923.
Gottschling, M. & Hilger, H.H. (2004): Characterization of a novel fruit type found in Ehretia (Ehretiaceae, Boraginales). - Blumea 49: 145-153.
Gottschling, M. (2004): Floral ontogeny in Bourreria (Ehretiaceae, Boraginales). - Flora 199 (5): 409-423.
Gottschling, M. & Hilger, H.H.(2003): First fossil record of transfer cells in angiosperms. - Amer. J. Bot. 90 (6): 957-959.
Gottschling, M., Mai, D.H. & Hilger, H.H. (2002): The systematic position of Ehretia fossils (Ehretiaceae,Boraginales) from the European Tertiary and implications for character evolution. - Review of Palaeobotany and Palynology 121 (2): 149-156.
Gottschling, M. & Hilger, H.H. (2001): Phylogenetic analysis and character evolution of Ehretia and Bourreria (Ehretiaceae, Boraginales) and their allies based on ITS1 sequences. - Bot. Jahrb. Syst. 123: 249-268.

Codonaceae

Weigend, M. & Hilger, H.H. (2016): Codonaceae. pp. 137-140 in Kadereit, J.W. & Bittrich, V. (eds.): The Families and Genera of Vascular Plants Vol. 14: Flowering Plants. Eudicots. Aquifoliales, Boraginales, Bruniales, Dipsacales, Escalloniales, Garryales, Paracryphiales, Solanales (except Convolvulaceae), Icacinaceae, Metteniusaceae, Vahliaceae. Springer - Switzerland.
Jeiter, J.; Danisch, F.& Hilger, H.H. (2016): Polymery and nectary chambers in Codon (Codonaceae) - flower and fruit development in a small, capsule-bearing family of Boraginales. - Flora 220: 94-102.
Weigend, M. & Hilger, H.H. (2010): Codonaceae - a newly required family name in Boraginales. - Phytotaxa 10: 26-30.

Wellstediaceae

Hilger, H.H. & Weigend, M. (2016): Wellstediaceae pp. 403-406 in Kadereit, J.W. & Bittrich, V. (eds.): The Families and Genera of Vascular Plants Vol. 14: Flowering Plants. Eudicots. Aquifoliales, Boraginales, Bruniales, Dipsacales, Escalloniales, Garryales, Paracryphiales, Solanales (except Convolvulaceae), Icacinaceae, Metteniusaceae, Vahliaceae. Springer - Switzerland.

Heliotropiaceae

Diane, N., Hilger, H.H., Förther, H., Weigend, M. & Luebert, F. (2016): Heliotropiaceae. pp. 203-211 in Kadereit, J.W. & Bittrich, V. (eds.): The Families and Genera of Vascular Plants Vol. 14: Flowering Plants. Eudicots. Aquifoliales, Boraginales, Bruniales, Dipsacales, Escalloniales, Garryales, Paracryphiales, Solanales (except Convolvulaceae), Icacinaceae, Metteniusaceae, Vahliaceae. Springer - Switzerland.
Luebert, F. & H.H. Hilger (2014): Typification of Heliotropium and Tournefortia (Heliotropiaceae) species described by Ruiz and Pavón. - AJB Madrid 71 (2) epub doi: http://dx.doi.org/10.3989/ajbm.2332.
Luebert, F.; Jacobs, P.; Hilger, H.H. & Muller, L.A. (2013 online): Evidence for non-allopatric speciation among closely related sympatric Heliotropium species in the Atacama Desert. - Ecol. Evol. online: http://onlinelibrary.wiley.com/doi/10.1002/ece3.929/full.
Luebert, F. (2013): A revision of Heliotropium Sect. Cochranea (Heliotropiaceae) - Kew Bulletin 68(1): 1–54.
Luebert, F. & Weigend, M. (2012): Three new species of Heliotropium sect. Heliothamnus (Heliotropiaceae) from Peru. - Phytotaxa 49: 35–44.
Luebert, F.; Hilger, H.H. & Weigend, M. (2011): Diversification in the Andes: Age and origins of South American Heliotropium lineages (Heliotropiaceae, Boraginales). - Molec. Phylogenet. Evol. 61 (1): 90-102.
Luebert, F.; Brokamp, G.; Wen, J.; Weigend, M. & Hilger, H.H. (2011): Phylogenetic relationships and morphological diversity in Neotropical Heliotropium (Heliotropiaceae). - Taxon 60 (3): 663-680.
Ibáñez, S.; Luebert, F. & Gómez, M. (2011): Primer registro de Heliotropium amplexicaule (Heliotropiaceae) en Chile. Gayana Bot. 68 (1): 93-96.
Luebert, F.; Weigend, M. & Hilger, H.H. (2010): Epitypification of Heliotropium arborescens L. (Heliotropiaceae). - Taxon 59 (4): 1263-1266.
Luebert, F. & Wen, J. (2008): Phylogenetic analysis and evolutionary diversification of Heliotropium Sect. Cochranea (Heliotropiaceae) in the Atacama Desert. - Syst. Bot. 33 (2): 390-402.
Hilger, H.H. & Diane, N. (2003): A systematic analysis of Heliotropiaceae (Boraginales) based on trnL and ITS1 sequence data. - Bot. Jahrb. Syst. 125 (1): 19-51.
Diane, N., Jakob, C. & Hilger, H.H. (2003): Comparative leaf anatomy and distribution of foliar trichome types in Heliotropiaceae - Flora 198: 468-485.
Weigend, M., Brokamp, G., Kirbach, A. & Förther, H.(2003): Notas sobre Heliotropium krauseanum Fedde, la unica especie de Heliotropium Sect. Cochranea del Perú (Heliotropiaceae = Boraginaceae subfam. Heliotropioideae). - Arnaldoa 10(1): 61-74.
Diane, N., Förther, H. & Hilger, H.H. (2002): Molecular Systematics of the Heliotropiaceae (Boraginales) Inferred from ITS1 Sequences. - Amer. J. Bot. 89 (2): 287- 297.
Hilger, H.H. (1992): Morphology of Heliotropium (Boraginaceae) dispersal units. Botanica Acta 105: 387-393.
Hilger, H.H. (1989): Flower and fruit development in the Macaronesian endemic Ceballosia fruticosa (syn. Messerschmidia fruticosa, Boraginaceae, Heliotropioideae). - Plant Systematics and Evolution 166: 119-129.
Hilger, H.H. (1987): Fruchtbiologische Untersuchungen an Heliotropioideae (Boraginaceae). I. Die Ontogenie der monospermen Früchte von Heliotropium supinum L. - Flora 179: 291-303.

Hydrophyllaceae incl. Namaceae

Hofmann, M., Walden, G.K., Hilger, H.H. & Weigend, M. (2016): Hydrophyllaceae. pp. 221-238 in Kadereit, J.W. & Bittrich, V. (eds.): The Families and Genera of Vascular Plants Vol. 14: Flowering Plants. Eudicots. Aquifoliales, Boraginales, Bruniales, Dipsacales, Escalloniales, Garryales, Paracryphiales, Solanales (except Convolvulaceae), Icacinaceae, Metteniusaceae, Vahliaceae. Springer - Switzerland.
Hofmann, M. (1999): Flower and fruit development in the genus Phacelia (Phacelieae, Hydrophyllaceae): characters of systematic value. - Syst. Geogr. Pl. 68(1-2): 203-212.
Hilger, H.H. (1987): Flower and fruit development in Wigandia caracasana (Hydrophyllaceae). - American Journal of Botany 74: 250-259.